Hoary marmot behavior

Hoary marmots, unlike most other marmots, have a very loose social system. Some groups display a dominance hierarchy as such:

Adult males > adult females > 2 yr olds > yearlings > 1st year marmots.

I found it interesting that where there is plentiful food, hoary marmots will live in colonies. However, where there is not much food, colonies do not exist. When hoary marmots live in colonies males will mate with multiple females and fight with other males for dominance. However when hoary marmots do not live in colonies they will only mate with one female and there is limited conflict between males.